Travels in CSR space adventures with cellular automata
- Slides: 169
Travels in (C-S-R) space: adventures with cellular automata Presentation ready with acknowledgements to Ric Colasanti (Corvallis) Andrew Askew (Sheffield)
CA in a community of virtual plants Contrasting tones represent patches of
This is a single propagule of a virtual plant It is about to grow in a resource-rich above- and below-ground environment
The plant has produced abundant growth aboveand below-ground and zones of resource depletion have appeared
Above-ground binary tree ( = shoot system) Each plant is built-up like this A branching module Above-ground array Above-ground binary tree base module Below-ground array Below-ground binary tree base module This is only a diagram, not a painting ! An end module Below-ground binary tree ( = root system)
The end-modules capture resources: Light and carbon dioxide from aboveground Water and nutrients from belowground The branching modules (parent or offspring) can pass resources to any adjoining modules In this way whole plants can grow
The virtual plants interact with their environment (and with their neighbours) just like real ones do They possess most of the properties of real individuals and populations For example …
Size Time Partitioning between root and shoot S-shaped growth curves Allometric coefficient Individual size Foraging towards resources Below-ground resource Functional equilibria Selfthinning line Population density Self-thinning in crowded populations
All of these plants have the same specification (modular rulebase) And this specification can easily be changed if we want the plants to behave differently… For example, we can recreate J P Grime’s system of C-S-R plant functional types But what is that exactly?
‘ The external factors which limit the amount of living and dead plant material present in any habitat may be classified into two categories ’ Opening sentence from J P Grime’s 1979 book Plant Strategies and Vegetation Processes
Category 1: Stress Phenomena which restrict plant production e. g. shortages of light, water, mineral nutrients, or non-optimal temperature
Category 2: Disturbance Phenomena which destroy plant production e. g. herbivory, pathogenicity, trampling, mowing, ploughing, wind damage, frosting, droughting, soil erosion, burning
Habitats may experience stress and disturbance to any degree and in any combination Stress Disturbance
Low or moderate combinations of stress and disturbance can support vegetation … Stress Disturbance … but extreme combinations of stress and disturbance cannot
There are other ways of describing stress and disturbance Stress Habitat productivity (= resource level) Disturbance Habitat duration
In the domain where vegetation is possible … S Stress-tolerator where S is high but D is low Stress Competitor where both S and D are low C R Disturbance Ruderal where S is low but D is high … plant life has evolved different strategies for dealing with the different combinations
S So this is ‘C-S -R space’ … C R … and these are the ‘habitats’ where no plant life occurs at all
To navigate in C-S-R space we bend the universe a little … S C R
S C R
S C R
S C R
S C R
S C R
S C R
C R S
C S R
C S R
C R S
C R S
C R S
… and recognize an intermediate type C CSR R S
… with further intermediates here C CR R CS CSR SR S
… and yet more intermediates here CR R C CS CSR SR S
So, how does all this relate to real vegetation? The high dimensionality of real plant life is reduced to plant functional types “ There are many more actors on the stage than roles that can be played ”
And what does that mean, exactly? Functional types provide a continuous view of vegetation when relative abundances, and even identities, of constituent species are in flux Tools that allocate C-S-R type to species, and C-S-R position to whole communities, can link separate vegetation into one conceptual framework Then effects of environment or management on biodiversity, vulnerability and stability can be evaluated on a common basis
We can recreate C-S-R plant functional types within the selfassembling model … … if we change the rulebases controlling morphology, physiology and reproductive behaviour …
With three levels possible in each of three traits, 27 simple functional types could be constructed However, we model only 7 types; the other 20 would include Darwinian Demons that do not respect evolutionary tradeoffs
Let’s see some competition between different types of plant Initially we will use only two types …
Small size, rapid growth and fast reproduction Medium size, moderately fast in growth and reproduction
(Red enters its 2 nd generation)
White has won !
Now let’s see if white always wins This time, the opposition is rather different …
Medium size, moderately fast in growth and reproduction Large size, very fast growing, slow reproductio n
The huge blue type has outcompeted both of the white plants, both above- and belowground And the simulation has run out of space …
So competition can be demonstrated realistically … … but most real communities involve more than two types of plant
We need seven functional types to cover the entire range of variation shown by herbaceous plant life To a first approximation, these seven types can simulate complex community processes very realistically
For example, an equal mixture of all seven types can be grown together … … in an environment which has high levels of resource, both above- and below-ground
The blue type has eliminated almost everything except white and green types And the simulation has almost run out of space again …
Now let’s grow the equal mixture of all seven types again … … but this time the environment has low levels of mineral nutrient resource (as indicated by the many grey cells)
(a gap has appeared here)
(red tries to colonize)
(but is unsuccessful)
White, green and yellow finally predominate … … blue is nowhere to be seen … … and total biomass is much reduced
Environmental gradients can be simulated by increasing resource levels in steps Whittaker-type niches then appear for contrasting plant types within these gradients
(types)
Let’s grow the equal mixture of all seven types again … … but this time under an environmental gradient of increasing mineral nutrient resource
Greatest biodiversity is at intermediate stress
Remember that environmental disturbance was defined as ‘removal of biomass after it has been created’ Trampling is therefore a disturbance It can be simulated by removing shoot material from certain sizes of patch at certain intervals of time and in a certain number of places
So we grow the equal mixture of all seven types again … … under an environmental gradient of increasing ‘trampling’ disturbance
Greatest biodiversity is at intermediate disturbance … … but the final number of types is low
Environmental stress and disturbance can, of course, be applied together … … and this can be done in all forms and combinations
So, again we grow the equal mixture of all seven types … … but in all factorial combinations of seven levels of stress and seven levels of disturbance
Greatest biodiversity is at intermediate productivity
The biomass-driven ‘humpbacked’ relationship is one of the highest-level properties that real plant communities possess Yet it emerges from the model solely because of the resource-capturing activity of modules in the selfassembling plants
These are all real experiments with virtual plants … and the plant, population and community processes all emerge from the one modular rulebase We can now ‘plant’ whole communities of any kind and subject them to different environments or management regimes Then we can look at topics such as biodiversity, vulnerability, resistance, resilience, stability, habitat / community heterogeneity, etc.
And as the modular rulebase is simply a string of numbers 2314232122133 1 2 controls 3 which how big, how much, how long, how often … (seems familiar? ) … we can modify this 2314232122133 virtual genome 123 wherever we like either accurately 2 3 1 4 2 3 2 1 2 3 123 or inaccurately 2 3 1 4 2 3 2 1 2 2 3 2 1 123 and then follow the downstream consequences of GM
In real experiments with virtual plants … One overnight run on one PC Approx. 100 personyears of growth experiments (not including the transgenic work!) Any takers? http: //www. ex. ac. uk/~rh 2 03/
- Cellular automata matlab
- Csr space
- Heat travels through empty space by
- A small, solid particle that travels through space.
- It is a form of energy that travels through space
- Unscented trajectory chapter 5
- Joint space vs cartesian space
- World space computer
- Space junk the space age began
- Camera space to world space
- Huckleberry finn questions
- Duchess in alice in wonderland
- The adventures of huckleberry finn chapter 1
- Pristine adventures
- Is huck finn a bildungsroman
- N adventures
- Dramatic irony in the adventures of tom sawyer
- Adventures of tom sawyer
- Irony in the adventures of tom sawyer
- Lying in a pool of tears
- Px adventures
- Smartie the penguin
- Huck finn controversy
- Huck finn chapter 12
- Rhyming scheme meaning
- Tours by tolano
- Windsor castle
- Arabian adventures abu dhabi
- Realism in the adventures of huckleberry finn
- What is the climax of huckleberry finn
- Adventures are the pits
- Miami-dade county parks and recreation staff
- The adventures of odysseus summary
- What are the 12 adventures of odysseus
- The adventures of jasper woodbury
- Assembly language adventures
- The merry adventures of robin hood characters
- Disney learning adventures theme song
- Ogden nash isabel
- Al's relativistic adventures
- Enriqueta ramirez
- Smartie the penguin
- Phasels wood activity centre
- Literary terms epic
- Adventures of huckleberry finn themes
- Huckleberry finn map
- Huckleberry finn chapter 1
- Adventures in axway
- Tolano adventures
- Irony in huckleberry finn
- Chapter 8 mother to mother
- Adventures of an it leader
- Hvad viser overskudsgraden
- Csr kursus
- Lars moratis
- Saint gobain uk csr
- Moral principles of csr
- Ackerman's model of social responsibility
- Diss jvs user manual
- Clsparse
- Csr piramis
- Csr dalam etika bisnis
- Csr performance indicators
- Corporate social responsibility presentation
- Unido csr
- Driving forces of csr
- Coporate social responsibility
- Saint gobain uk csr
- Pandangan praktis realistis
- Csr arguments
- Csr architecture
- Csr reading strategy
- Archie carroll pyramid
- Filantrópikus
- Csr management system
- Csr arguments
- What is trusteeship management
- High latitude range management training alaska
- Introduction of csr
- Chief csr officer
- Csd csr
- Objectives of csr
- Amway csr activities
- Apa itu csr
- Chapter 4 ethics and social responsibility
- Saint gobain uk csr
- "raport eb"
- Nature of csr
- Marriott corporate responsibility
- Dějiny udatného českého národa vznik čsr
- Unido model of csr
- New model of csr
- Csr communication strategy
- Corporate social responsibility cos è
- Conscious marketing vs corporate social responsibility
- Csr full form
- Howard bowen corporate social responsibility
- Definition of csr
- Indikator keberhasilan csr
- Csr gdansk
- Ranjan mohapatra csr vision
- Five driving forces of csr
- Pertanyaan tanggung jawab sosial
- Selbstauskunftsfragebogen csr
- Bluetooth gaia
- Transcelerate csr template
- Concepts of csr
- Csr voluntary
- C s r is
- Contoh sk kp-spams pamsimas
- Csr formula
- Economic responsibility
- Archie carroll pyramid
- Aldi social responsibility
- Csr vznik
- Class system
- Interessentkart
- Confedency
- Kind of heat transfer
- Acceleration word problems
- Ehat is sound
- Light travels 3 00 000 km/s. is it velocity or speed
- What two measurements are necessary for calculating speed
- How is sound produced
- Examples of a medium in waves
- Diction in there will come soft rains
- Supaa travels
- Satire in gulliver's travels part 3
- Light always travels in a
- Abraham's journey map
- Gullivers travels quiz
- Example of mechanical wave
- A wave that travels along the surface separating two media
- Define lilliputian
- A snowmobile travels in a straight line
- Information travels between components on the motherboard
- Gulliver's travels quiz
- Grnconnect
- Which phrase correctly describes the tang dynasty?
- Odysseus travels
- Picture how light travels
- Electromatic goanimate
- What is the curved path traveled by the projectile
- Sound energy travels on a wave
- A car travels from point a to b in 3 hours
- Phantom tours and travels pvt ltd
- What is average speed
- How does gulliver earn the title of nardac in lilliput?
- Methods of heat transfer
- Gulliver's travels satire
- Viking trade routes
- "residual income"
- Diagram showing that light travels in a straight line
- Sound travels fastest in
- Appitravels
- Distance vs displacement
- A woman travels in a lift. the mass of the woman is 50 kg
- Chapter 17 mechanical waves and sound
- Satire in gulliver travels part 1
- Distance an object travels
- Enquiry letter for admission in college
- Wave speed equation
- What is pitch in sound
- Travels with charley themes
- 3 ways heat travels
- Gulliver's travels essay questions
- A go cart travels around a flat horizontal circular track
- Which travels along a surface separating two media
- Laputa castle in the sky
- How light travels diagram