1. What are trace fossils? Sedimentary structures produced by life - Burrows, borings, tracks, trails, roots. . .
. . . and dung! The Lloyds Bank coprolite
Why they matter • • • Fossil record of behaviour Fossil record of soft-bodied organisms Organism-environment interactions Ecological niche creation Nutrient flux Taphonomy Luminous lugworm
Uniformitarianism The present is the key to the past
Ichnological principles 1. Same organism; different traces Arthropod trace fossils
Ichnological principles 2. Different organisms; same traces Annelid worms Sea anemones Phoronids
Ichnological principles 3. Same trace; different preservation
2. Traces of Yorkshire What trace fossils does the museum have?
3. Common trace fossils (Burrows, trails; mostly marine)
Ichnotaxonomy N. B. Names are for trace fossil only, not organism that made the structure
Planolites
Skolithos
Thalassinoides
Thalassinoides tracemaker?
Chondrites
Ophiomorpha
Ophiomorpha tracemaker?
Rhizocorallium
Rusophycus
Rusophycus tracemaker?
Trace fossil? Impression of a jellyfish? Or a sand volcano?
Not trace fossils Shrinkage cracks
Not trace fossils Flute casts
Hand Specimens: What Have You Got?
4. The Fossil Record of Behaviour
Earliest animals? 565 Ma locomotion trails, Newfoundland: Liu et al. (2010) http: //geology. gsapubs. org/ content/38/2/123. abstract
Ediacaran-Cambrian boundary
Cambrian complexity
The Cambrian Explosion Seilacher & Pfluger (1994)
Onshore to offshore
Colonization of land
Plant Trace Fossils
Becoming upright Trackway analysis shows tetrapods become bipedal in early Triassic Kubo & Benton (2009) http: //onlinelibrary. wiley. com/doi/10. 1111 /j. 1475 -4983. 2009. 00897. x/full
The speed of dinosaurs
5. The speed of dinosaurs? • • • Type of footprint? Footprint size? Animal size? Stride length? Speed? Behaviour?
Calculating trackmaker speed v = 0. 25*g 0. 5*SL 1. 67*h-1. 17 Or, more simply: Hip height = 4 x footprint length Stride length/ hip height = speed (<2 = walking, >3 = running) Sheffield Geology Group – dinosaur speed calculator: http: //www. sorbygeology. group. shef. ac. uk/DINOC 01/dinocal 1. htm l