E coli Hfr H Hfr 1 Hfr 2

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六、E. coli 环状染色体 Hfr H Hfr 1 Hfr 2 Hfr 3 thr pro lac

六、E. coli 环状染色体 Hfr H Hfr 1 Hfr 2 Hfr 3 thr pro lac pur gal his gly thi thr thi gly his gal pur lac pro thr pro thi gly his gal pur lac pro thr thi gly his gal

4 七. 重组频率的计算 (一) 等级传递绘图 1956年Wollman, Jacob和 Hayes Hfr. H thr+ leu+ azis lac+

4 七. 重组频率的计算 (一) 等级传递绘图 1956年Wollman, Jacob和 Hayes Hfr. H thr+ leu+ azis lac+ gal+ (λ)+ str+ mal+ mtl+ F－ thr- leu- azir lac- gal - (λ)- str- mal- mtl-

(二) 中断杂交作图 (interrupted matingexperiment) 4 1961年 Jacob 和 Wollman 供体：Hfr H strs thr＋leu＋azir tonr

(二) 中断杂交作图 (interrupted matingexperiment) 4 1961年 Jacob 和 Wollman 供体：Hfr H strs thr＋leu＋azir tonr lac＋ gal＋ 受体：F－ strr thr leu azis tons lac－gal－

三. 共转导（contransduc）

三. 共转导（contransduc）

4 附着位点（attachment site , att ） att B（又叫attλ）， att P 4 正常的切离 4 非正常切离

4 附着位点（attachment site , att ） att B（又叫attλ）， att P 4 正常的切离 4 非正常切离 4 λdg(λd gal) 4 λpgal 4 λdbio 4 低频转导(low-frequency transduction, LFT） 4 高频转导(High-feequency transduction, HFT) 4 双重溶原（double lysoge）细菌

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